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Why do female adders copulate so frequently?Madsen, Thomas; Shine, Richard; Loman, Jon; Håkansson, Thomas · 1992
Determinants of mating success in male adders, Vipera berusThomas Madsen; Richard Shine; Jon Loman; Thomas Håkansson · 1993
A Better Title: "Female Benefits from Multiple Matings: A Model"Jon Loman, Thomas Madsen and Thomas Håkansson · Oikos · 1989
Reproductive Tactics of Large and Small Male Toads Bufo BufoLoman, Jon; Madsen, Thomas · Oikos · 1986
Increased Fitness from Multiple Matings, and Genetic Heterogeneity: A Model of a Possible MechanismJon Loman, Thomas Madsen and Thomas Håkansson · Oikos · 1988
Sex ratio of breeding Common toads (Bufo bufo) – influence of survival and skipped breedingLoman, Jon; Madsen, Thomas · Amphibia-Reptilia · 2010